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''Epidendrum secundum'', one of the crucifix orchids, is a poorly understood reed stemmed species, which Dressler (1989) describes as "the ''Epidendrum secundum'' complex." According to Dressler, there are dozens of varieties, some of which appear to deserve species rank. Arditti and Ghani note that ''E. secundum ''has the distinction of bearing the longest seeds known in the Orchidaceae, 6.0 mm long. By comparison, the seeds of ''E. ibaguense'' (another crucifix orchid) are only 2.9 mm long. Like the other members of ''E.'' subg. ''Amphiglottium'', ''E. secundum'' is a sympodial plant which has thin stems covered from the base with imbricating sheaths which are leaf bearing above, a terminal inflorescence covered at its base with thin imbricating sheaths, and flowers with the lip adnate to the column to its apex. The flowers are non-resupinate (unlike ''E. ibaguense'' and ''E. radicans''), can come in shades of lilac, red, orange, or yellow, and feature a notable callus on the fringed trilobate lip. The plant is rather cool-growing and can tolerate a light frost. == Diversity within the complex == In 1861, H. R. Reichenbach presented a treatment of the genus ''Epidendrum'', including his view of the sub-section ''Tuberculata''. Correlating his list of (in Reichenbach's view) separate species with the Kew Monocot Checklist (July 16, 2009) yielded the following list of taxa "which appear to deserve species rank" that Kew checklist reviewers consider to be synonyms for ''E. secundum'' (the page numbers refer to Reichenbach 1861): * ''E. ansiferum'' Rchb.f. & Warsz., Bonplandia (Hannover) 2: 111 (1854). (pp 394-395) * ''E. brachyphyllum'' Lindl., Fol. Orchid. 3:72(1853) (p. 392) * ''E. fimbria'' Rchb.f., Bonplandia (Hannover) 2: 282 (1854). (p. 394) * ''E. gracilicaule'' Rchb.f. & Warsz., Bonplandia (Hannover) 2: 111 (1854) (p. 392) * ''E. incisum'' Rchb.f. & Warsz., Bonplandia (Hannover) 2: 112 (1854), nom. illeg. (p.394) * ''E. lacerum'' Lindl., Edwards's Bot. Reg. 24(Misc.): 17 (1838). (pp. 395-396) * ''E. lindenii'' Lindl., Edwards's Bot. Reg. 31(Misc.): 48 (1845), nom. illeg (p. 393) * ''E. novogranatense'' Rchb.f. & Warsz., Bonplandia (Hannover) 2: 111 (1854). (p. 396) The diversity of ''E. secundum'' is further demonstrated by counts of the chromosome number (in root tissues): a lilac-flowered individual from Bolivia was found to have 2''n'' = 28; a lilac-flowered individual from Santo Antônio do Itambé, Brazil, 2''n'' = 52; an orange-flowered individual from Santo Antônio do Itambé, Brazil, 2''n'' = 48; a lilac-flowered individual from Serra do Rio do Rastro, Brazil, 2''n'' = 40; a lilac-flowered individual from Venezuela, 2''n'' = 80; all in disagreement with a previously reported value of 2''n'' = 68. The Kew Monocot checklist lists many more binomials as synonyms of ''E. secundum'':〔http://apps.kew.org/wcsp/synonomy.do?accepted_id=69534&repSynonym_id=-9998&name_id=69534&status=true〕 * ''E. antioquiense'' Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 125 (1920) * ''E. bulkeleyi'' A.D.Hawkes, Orquídea (Rio de Janeiro) 18: 168 (1957) * ''E. coroicoense'' Schltr., Repert. Spec. Nov. Regni Veg. 27: 60 (1929) * ''E. corymbosum'' Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil.: 246 (1798) * ''E. corymbosum'' var. ''latifolium'' Cogn. in C.F.P.von Martius & auct. suc. (eds.), Fl. Bras. 3(5): 145 (1898) * ''E. crassifolium'' var. ''albescens'' Pabst, Bradea 2: 64 (1976) * ''E. cuzcoense'' Schltr., Repert. Spec. Nov. Regni Veg. Beih. 9: 82 (1921) * ''E. dolichopus'' Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 131 (1920) * ''E. elongatum'' Jacq., Collectanea 3: 260 (1789) * ''E. fastigiatum'' Lindl., Fol. Orchid. 3: 71 (1853) * ''E. giroudianum'' Rchb.f., Bonplandia (Hannover) 4: 327 (1856) * ''E. herzogii'' Schltr., Repert. Spec. Nov. Regni Veg. 12: 489 (1913) * ''E. inconstans'' Ames ex Gleason, Bull. Torrey Bot. Club 58: 350 (1931) * ''E. lacera'' (Lindl.) Britton, Sci. Surv. Porto Rico & Virgin Islands 5: 201 (1924) * ''E. longihastatum'' Barb.Rodr., Gen. Spec. Orchid. 1: 59 (1877) * ''E. pachyphyllum'' Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 140 (1920) * ''E. polyschistum'' Schltr., Repert. Spec. Nov. Regni Veg. Beih. 7: 143 (1920) * ''E. secundum'' var. ''albescens'' (Pabst) F.Barros, Acta Bot. Brasil. 10: 142 (1996) * ''E. secundum'' f. ''albescens'' (Pabst) F.Barros, Hoehnea 29: 111 (2002) * ''E. sulfuratorium'' E.H.L.Krause, Beih. Bot. Centralbl. 36(2): 336 (1914) * ''E. tarmense'' Schltr., Repert. Spec. Nov. Regni Veg. Beih. 9: 94 (1921) * ''E. tricallosum'' Schltr., Repert. Spec. Nov. Regni Veg. Beih. 6: 39 (1919) * ''E. versicolor'' Hoehne & Schltr., Arch. Bot. São Paulo 1: 245 (1926) 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Epidendrum secundum」の詳細全文を読む スポンサード リンク
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